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Du Toit went far beyond Wegener in his understanding of Gondwanaland; he was able to reconstruct both the early merging of the various continental pieces and their The End of Evolution climactic melding into a single continental mass in Late Paleozoic time, followed by their subsequent fragmentation during the Mesozoic and Cenozoic eras. He became versed in the stratigraphy not only of his own continent, but of the other Gondwana members as well. Perhaps his most telling argument supporting continental drift was his demonstration of the remarkable similarity in Late Paleozoic rock sequences on the various continental pieces.

But catastrophic as that great extinction was, it too slowed life's relentless expansion only momentarily. Following the Second Event mammals began their hegemony, in tandem with flowering plants, insects, mollusks, and an entirely new suite of oceanic plankton. In the modern day, levels of diversity are far, far higher than at any time in the deep past. But the long history of life on the earth offers little congratulation: Many past empires of life stood tall as well, only to slide back into oblivion and dust.

Any mass extinction has two parts: mass death followed by the earth's repopulation with a largely new suite of creatures. Cuvier's proof of the reality of extinction solved the first part of the puzzle. But what of the repopulation? The origin of species remained an abominable mystery to most scientists of the early nineteenth century; many, such as the pioneering French geologist Alcide d'Orbigny, were content to leave both the extinction and repopulation to God. Others were not so sure. One such skeptic was Charles Darwin, whose great work gave Phillips the key to the second part of the puzzle: The repopulation of the earth occurred through speciation.

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